Will PCIe x1 x4 cards work in x16 slot?

What is the purpose of the PCIe X1 slot pcie x1 in x16 slot PCIe x16 slots are generally intended for use with Graphics Cards PCIe 1x slots are generally for smaller add-in cards like Wifi Adapters, USB Adapters, or pcie 3 slot Therefore, users can insert a card using PCIe x1 interface into a PCIe x4, x8 or x16 slots when the PCIe x1 slot has been occupied, which solves the problem of

pcie 4 in pcie 3 slot ssd A lot of budget motherboards, having a x16x8x4 and 3x x1 slots, actually don't handle over 4 GPUs Even if you somehow could run the first GPU